Emails getting returned to customers; "Sender address quota exceeded"

Some of my customers are reporting that their emails are being returned to them, with a "Sender address quota exceeded" message. When I login to the webmail interface (Zimbra / Homestead ... not sure what the right name is), it shows that I'm well within my quota. What is going on? Can this problem be rectified so that I don't miss any emails?? I'm still getting most of my emails I think, but at least 3 people have forwarded this to me on my other support address.
The returned email shows something like this:

Delivery to the following recipient failed permanently:

bugreport@zaal-llc.com

Technical details of permanent failure:
Google tried to deliver your message, but it was rejected by the server for the recipient domain zaal-llc.com by inbound.homesteadmail.com. [209.157.71.80].

The error that the other server returned was:
554 5.7.1 : Recipient address rejected: Policy Rejection- Sender address quota exceeded: Throttled (Error code ERR-3375-T).